LineageOS always uses multiple Google services while giving them privileged access even if users don't use microG. It would be wrong to imply they don't. GrapheneOS doesn't use Google services by default.

To clarify further for what they always use Google services for even without microG, they use them for:

Connectivity checks

Network time

Attestation key provisioning

SUPL

DNS fallback

PSDS (Pixel 6 and 7)

eSIM activation

and more enabled by default.